:root {
  --green: #0a963d;
  --green-dark: #075d2c;
  --green-deep: #063d24;
  --green-pale: #edf6ef;
  --ink: #172019;
  --muted: #68736b;
  --line: rgba(13, 72, 37, .14);
  --paper: #fff;
  --canvas: #f4f7f4;
  --header-height: 76px;
  --shadow: 0 24px 70px rgba(20, 57, 35, .12);
}

* { box-sizing: border-box; }
html { scroll-behavior: smooth; scroll-padding-top: var(--header-height); }
body {
  margin: 0;
  background: var(--paper);
  color: var(--ink);
  font-family: Arial, Helvetica, sans-serif;
  line-height: 1.55;
  overflow-x: hidden;
}
a { color: inherit; text-decoration: none; }
img { display: block; max-width: 100%; }
button, input, select, textarea { font: inherit; }

.sr-only {
  position: absolute;
  width: 1px;
  height: 1px;
  padding: 0;
  margin: -1px;
  overflow: hidden;
  clip: rect(0, 0, 0, 0);
  white-space: nowrap;
  border: 0;
}

.scroll-progress {
  position: fixed;
  z-index: 60;
  top: 0;
  right: 0;
  left: 0;
  height: 3px;
  pointer-events: none;
}
.scroll-progress span {
  display: block;
  width: 100%;
  height: 100%;
  background: var(--green);
  transform: scaleX(0);
  transform-origin: left center;
}

.site-header {
  position: sticky;
  z-index: 50;
  top: 0;
  display: flex;
  align-items: center;
  justify-content: space-between;
  min-height: var(--header-height);
  padding: 10px max(28px, calc((100vw - 1240px) / 2));
  background: rgba(255, 255, 255, .88);
  border-bottom: 1px solid rgba(5, 62, 30, .08);
  backdrop-filter: blur(18px);
}
.brand img { width: 132px; height: auto; }
.desktop-nav { display: flex; align-items: center; gap: 28px; }
.desktop-nav > a,
.nav-dropdown > a {
  position: relative;
  padding: 12px 0;
  color: #4f5b53;
  font-size: 15px;
  font-weight: 700;
}
.desktop-nav > a::after,
.nav-dropdown > a::after {
  position: absolute;
  right: 0;
  bottom: 5px;
  left: 0;
  height: 2px;
  background: var(--green);
  content: '';
  transform: scaleX(0);
  transform-origin: right;
  transition: transform .25s ease;
}
.desktop-nav > a:hover::after,
.desktop-nav > a:focus-visible::after,
.desktop-nav a.active::after,
.nav-dropdown > a:hover::after,
.nav-dropdown > a:focus-visible::after,
.nav-dropdown > a.active::after { transform: scaleX(1); transform-origin: left; }
.desktop-nav a.active { color: var(--green-dark); }
.nav-dropdown { position: relative; }
.dropdown-panel {
  position: absolute;
  top: 48px;
  left: -22px;
  display: grid;
  min-width: 165px;
  padding: 9px;
  visibility: hidden;
  background: rgba(255, 255, 255, .98);
  border: 1px solid var(--line);
  border-radius: 16px;
  box-shadow: var(--shadow);
  opacity: 0;
  transform: translateY(8px);
  transition: opacity .2s ease, transform .2s ease, visibility .2s;
}
.nav-dropdown:hover .dropdown-panel,
.nav-dropdown:focus-within .dropdown-panel { visibility: visible; opacity: 1; transform: translateY(0); }
.dropdown-panel a { padding: 11px 13px; border-radius: 10px; color: #4c5850; font-weight: 700; }
.dropdown-panel a:hover,
.dropdown-panel a:focus-visible { background: var(--green-pale); color: var(--green-dark); }

.language-switch {
  display: inline-flex;
  align-items: center;
  gap: 5px;
  padding: 7px 11px;
  border-radius: 999px;
  background: var(--green-pale);
  color: #839188;
  font-size: 11px;
  font-weight: 800;
}
.language-switch i { color: #a9b5ad; font-style: normal; }
.language-switch a.active { color: var(--green-dark); }

.mobile-nav { display: none; position: relative; }
.mobile-nav summary { display: grid; gap: 5px; padding: 10px; cursor: pointer; list-style: none; }
.mobile-nav summary::-webkit-details-marker { display: none; }
.mobile-nav summary span { display: block; width: 25px; height: 2px; background: var(--ink); }
.mobile-nav nav {
  position: absolute;
  top: 46px;
  right: 0;
  display: grid;
  min-width: 220px;
  padding: 12px;
  background: #fff;
  border: 1px solid var(--line);
  border-radius: 18px;
  box-shadow: var(--shadow);
}
.mobile-nav nav > a { padding: 11px 13px; border-radius: 10px; font-weight: 700; }
.mobile-nav nav > a:hover { background: var(--green-pale); }
.mobile-language { display: flex; gap: 8px; margin-top: 8px; }
.mobile-language a { flex: 1; padding: 8px; border-radius: 999px; background: var(--green-pale); color: var(--green-dark); text-align: center; font-size: 12px; font-weight: 800; }

.page-anchor { scroll-margin-top: var(--header-height); }
.hero-section {
  position: relative;
  display: grid;
  min-height: calc(100svh - var(--header-height));
  place-items: center;
  overflow: hidden;
  padding: 88px 28px 100px;
  background:
    radial-gradient(circle at 78% 18%, rgba(78, 190, 116, .22), transparent 31%),
    radial-gradient(circle at 9% 75%, rgba(208, 232, 213, .8), transparent 34%),
    linear-gradient(145deg, #f9fcf9 0%, #eef7f0 58%, #fff 100%);
}
.hero-section::before,
.hero-section::after {
  position: absolute;
  border: 1px solid rgba(10, 150, 61, .12);
  border-radius: 50%;
  content: '';
}
.hero-section::before { width: 520px; height: 520px; top: -310px; right: -100px; }
.hero-section::after { width: 390px; height: 390px; bottom: -260px; left: -90px; }
.hero-glow {
  position: absolute;
  width: min(72vw, 920px);
  height: min(72vw, 920px);
  border-radius: 50%;
  background: radial-gradient(circle, rgba(15, 155, 65, .13), transparent 66%);
  animation: breathe 7s ease-in-out infinite;
}
@keyframes breathe { 50% { transform: scale(1.12); opacity: .72; } }
.hero-content { position: relative; z-index: 2; width: min(980px, 100%); text-align: center; }
.eyebrow {
  margin: 0 0 16px;
  color: var(--green-dark);
  font-size: 12px;
  font-weight: 800;
  letter-spacing: .16em;
  text-transform: uppercase;
}
.hero-logo-frame {
  width: min(900px, 100%);
  margin: 0 auto;
  padding: clamp(20px, 4vw, 42px);
  background: rgba(255, 255, 255, .78);
  border: 1px solid rgba(255, 255, 255, .9);
  box-shadow: 0 35px 100px rgba(16, 77, 38, .12);
  backdrop-filter: blur(12px);
  transition: transform .2s linear;
}
.hero-logo-frame img { width: 100%; height: auto; }
.hero-lead { max-width: 760px; margin: 35px auto 0; color: #34473a; font-size: clamp(17px, 2vw, 21px); line-height: 1.65; }
.hero-actions { display: flex; align-items: center; justify-content: center; gap: 24px; margin-top: 34px; }
.primary-button,
.dataset-button {
  display: inline-flex;
  align-items: center;
  justify-content: center;
  gap: 12px;
  min-height: 48px;
  padding: 12px 22px;
  border: 1px solid var(--green-dark);
  border-radius: 999px;
  background: var(--green-dark);
  color: #fff;
  font-weight: 800;
  cursor: pointer;
  transition: transform .2s ease, background .2s ease, color .2s ease, box-shadow .2s ease;
}
.primary-button:hover,
.primary-button:focus-visible { background: #fff; color: var(--green-dark); box-shadow: 0 12px 32px rgba(7, 93, 44, .16); transform: translateY(-2px); }
.text-link { padding: 11px 0; color: var(--green-dark); font-weight: 800; }
.text-link span { display: inline-block; margin-left: 7px; transition: transform .2s ease; }
.text-link:hover span { transform: translateY(4px); }
.scroll-cue { position: absolute; z-index: 2; bottom: 25px; width: 28px; height: 46px; border: 1px solid rgba(7, 93, 44, .4); border-radius: 999px; }
.scroll-cue span { position: absolute; top: 9px; left: 50%; width: 4px; height: 8px; border-radius: 999px; background: var(--green-dark); transform: translateX(-50%); animation: scroll-cue 1.8s ease infinite; }
@keyframes scroll-cue { 70% { transform: translate(-50%, 15px); opacity: 0; } 71% { transform: translate(-50%, 0); opacity: 0; } }

.page-section { position: relative; padding: clamp(88px, 10vw, 140px) 28px; }
.section-shell { position: relative; z-index: 2; width: min(1180px, 100%); margin: 0 auto; }
.section-heading { max-width: 800px; margin-bottom: 58px; }
.section-heading h2,
.contact-copy h2 { margin: 0; font-size: clamp(34px, 4.4vw, 52px); line-height: 1.08; letter-spacing: -.035em; }
.section-heading > p:last-child,
.contact-copy > p { max-width: 720px; margin: 22px 0 0; color: var(--muted); font-size: 18px; line-height: 1.75; }

.about-section { overflow: hidden; background: var(--canvas); }
.about-section::before {
  position: absolute;
  top: 120px;
  right: -140px;
  width: 630px;
  height: 220px;
  background: url('../images/cromocat-hero.jpg') center / contain no-repeat;
  content: '';
  filter: grayscale(1);
  opacity: .035;
  transform: rotate(-8deg);
}
.about-grid { display: grid; grid-template-columns: .75fr 1.25fr; gap: 22px; }
.glass-card {
  position: relative;
  min-height: 260px;
  padding: 34px;
  overflow: hidden;
  background: rgba(255, 255, 255, .82);
  border: 1px solid rgba(255, 255, 255, .9);
  box-shadow: 0 18px 54px rgba(20, 57, 35, .07);
}
.glass-card::after { position: absolute; right: -45px; bottom: -55px; width: 150px; height: 150px; border-radius: 50%; background: var(--green-pale); content: ''; }
.card-index { color: var(--green); font-size: 12px; font-weight: 900; letter-spacing: .16em; }
.glass-card h3,
.related-panel h3,
.form-card h3 { margin: 22px 0 18px; font-size: 25px; }
.clean-list,
.link-list { position: relative; z-index: 1; margin: 0; padding-left: 20px; line-height: 1.9; }
.link-list a,
.publication-list a,
.citation-panel a,
.email-link { color: var(--green-dark); text-decoration: underline; text-underline-offset: 3px; }

.institutional-strip {
  display: grid;
  grid-template-columns: 140px minmax(220px, 285px) minmax(220px, 300px);
  align-items: center;
  justify-content: center;
  gap: clamp(35px, 7vw, 90px);
  margin: 60px 0;
  padding: 35px 0;
  border-top: 1px solid var(--line);
  border-bottom: 1px solid var(--line);
}
.institutional-strip > img:first-child { width: 140px; }
.institutional-strip > img:nth-child(2) { width: 285px; }
.institutional-strip a img { width: 290px; }

.about-secondary { display: grid; grid-template-columns: 1fr 1fr; gap: 22px; }
.related-panel,
.citation-panel { padding: 34px; background: #fff; border-top: 3px solid var(--green); box-shadow: 0 14px 45px rgba(20, 57, 35, .06); }
.related-panel > a { display: grid; grid-template-columns: 100px 1fr; gap: 15px; align-items: center; padding: 15px 0; border-top: 1px solid var(--line); }
.related-panel strong { color: var(--green-dark); }
.related-panel span,
.citation-panel { color: #526057; }
.citation-panel p:last-child { margin: 20px 0 0; line-height: 1.75; }

.about-details { margin-top: 28px; border-top: 1px solid var(--line); }
.about-details details { border-bottom: 1px solid var(--line); }
.about-details summary { display: flex; align-items: center; justify-content: space-between; padding: 24px 4px; cursor: pointer; list-style: none; color: var(--green-dark); font-size: 19px; font-weight: 800; }
.about-details summary::-webkit-details-marker { display: none; }
.about-details summary span { font-size: 26px; transition: transform .25s ease; }
.about-details details[open] summary span { transform: rotate(45deg); }
.publication-list { display: grid; gap: 17px; margin: 0 0 28px; padding-left: 27px; line-height: 1.7; }
.publication-list strong { display: inline-block; min-width: 54px; color: var(--green-dark); }
.collaborator-intro { max-width: 960px; margin: 0 0 22px; color: #46544b; line-height: 1.65; }
.name-cloud { display: flex; flex-wrap: wrap; gap: 9px; margin: 0 0 28px; padding: 0; list-style: none; }
.name-cloud li { padding: 8px 13px; border-radius: 999px; background: #fff; border: 1px solid var(--line); font-size: 14px; }
.funding-note { margin: 34px 0 0; color: var(--muted); font-size: 14px; }

.search-section { min-height: 84vh; overflow: hidden; background: linear-gradient(135deg, #063d24 0%, #075d2c 55%, #087e37 100%); color: #fff; }
.search-backdrop { position: absolute; inset: 0; background: url('../images/cromocat-hero.jpg') 92% 50% / min(850px, 70vw) auto no-repeat; filter: grayscale(1); mix-blend-mode: screen; opacity: .055; transform: scale(1.08); }
.section-heading--light .eyebrow { color: #bde9c9; }
.section-heading--light > p:last-child { color: #d4e8d9; }
.dataset-grid { display: grid; grid-template-columns: repeat(3, 1fr); gap: 18px; }
.dataset-card { display: flex; flex-direction: column; min-height: 430px; padding: 32px; background: rgba(255, 255, 255, .085); border: 1px solid rgba(255, 255, 255, .18); backdrop-filter: blur(12px); transition: transform .28s ease, background .28s ease, border-color .28s ease; }
.dataset-card:hover { background: rgba(255, 255, 255, .13); border-color: rgba(255, 255, 255, .34); transform: translateY(-8px); }
.dataset-number { color: #a9e3b8; font-size: 12px; font-weight: 900; letter-spacing: .16em; }
.dataset-card h3 { margin: 50px 0 20px; font-size: 30px; }
.dataset-card p { margin: 0 0 28px; color: #d8e8dc; line-height: 1.7; }
.dataset-button { align-self: flex-start; margin-top: auto; border-color: #fff; background: #fff; color: var(--green-dark); }
.dataset-button:hover { background: transparent; color: #fff; transform: translateY(-2px); }

.contact-section { background: #fff; }
.contact-shell { display: grid; grid-template-columns: .82fr 1.18fr; gap: clamp(45px, 8vw, 105px); align-items: start; }
.contact-copy { position: sticky; top: calc(var(--header-height) + 38px); }
.email-link { display: inline-block; margin-top: 25px; font-size: 17px; font-weight: 800; }
.form-card { padding: clamp(27px, 4vw, 45px); background: var(--canvas); border: 1px solid var(--line); box-shadow: var(--shadow); }
.form-card h3 { margin-top: 0; }
.contact-form { display: grid; gap: 13px; }
.contact-form label:not(.consent-row) { display: grid; gap: 7px; color: #334138; font-size: 13px; font-weight: 800; }
.contact-form label span { color: #7a857e; font-weight: 400; }
.field-row { display: grid; grid-template-columns: 1fr 1fr; gap: 13px; }
.contact-form input,
.contact-form select,
.contact-form textarea { width: 100%; padding: 12px 13px; border: 1px solid #cbd7cf; border-radius: 10px; background: #fff; color: var(--ink); }
.contact-form textarea { resize: vertical; }
.contact-form input:focus,
.contact-form select:focus,
.contact-form textarea:focus { outline: 3px solid rgba(10, 150, 61, .15); border-color: var(--green); }
.consent-row { display: grid; grid-template-columns: 20px 1fr; gap: 10px; align-items: start; margin: 8px 0; color: #59655d; font-size: 12px; }
.consent-row input { width: 17px; margin-top: 3px; }
.honeypot { position: absolute; left: -10000px; width: 1px; height: 1px; overflow: hidden; }
.form-status { margin: 0 0 20px; padding: 14px 16px; border-left: 3px solid; background: #fff; line-height: 1.55; }
.form-status.success { border-color: var(--green); }
.form-status.error { border-color: #b42318; }
.form-status p { margin: 0 0 8px; font-weight: 800; }
.form-status ul { margin: 0; padding-left: 20px; }

.site-footer { display: grid; grid-template-columns: 150px 1fr auto; align-items: center; gap: 30px; padding: 34px max(28px, calc((100vw - 1240px) / 2)); background: #061f14; color: #c5d6cb; font-size: 12px; }
.footer-brand {
  display: inline-flex;
  justify-self: start;
  width: max-content;
  padding: 9px 13px;
  background: #fff;
  border-radius: 10px;
  box-shadow: 0 6px 20px rgba(0, 0, 0, .14);
}
.footer-brand img { width: 142px; filter: none; }
.site-footer p { margin: 0; }
.site-footer nav { display: flex; gap: 18px; }
.site-footer a:hover { color: #fff; }

.js [data-reveal] { opacity: 0; transform: translateY(34px); transition: opacity .75s ease, transform .75s cubic-bezier(.2, .7, .2, 1); }
.js [data-reveal].is-visible { opacity: 1; transform: translateY(0); }
.js .dataset-card:nth-child(2) { transition-delay: .09s; }
.js .dataset-card:nth-child(3) { transition-delay: .18s; }

@media (max-width: 940px) {
  .desktop-nav { display: none; }
  .mobile-nav { display: block; }
  .about-grid,
  .about-secondary,
  .contact-shell { grid-template-columns: 1fr; }
  .institutional-strip { grid-template-columns: 125px minmax(200px, 260px); }
  .institutional-strip a { grid-column: 1 / -1; justify-self: center; }
  .dataset-grid { grid-template-columns: 1fr; }
  .dataset-card { min-height: 390px; }
  .dataset-card h3 { margin-top: 28px; }
  .contact-copy { position: static; }
  .site-footer { grid-template-columns: 130px 1fr; }
  .site-footer nav { grid-column: 1 / -1; }
}

@media (max-width: 620px) {
  :root { --header-height: 68px; }
  .site-header { padding-inline: 18px; }
  .brand img { width: 114px; }
  .hero-section { min-height: calc(100svh - var(--header-height)); padding: 70px 18px 90px; }
  .hero-logo-frame { padding: 18px; }
  .hero-lead { font-size: 16px; }
  .hero-actions { flex-direction: column; gap: 8px; }
  .scroll-cue { display: none; }
  .page-section { padding: 80px 18px; }
  .section-heading { margin-bottom: 40px; }
  .section-heading h2,
  .contact-copy h2 { font-size: 34px; }
  .section-heading > p:last-child,
  .contact-copy > p { font-size: 16px; }
  .glass-card,
  .related-panel,
  .citation-panel,
  .dataset-card,
  .form-card { padding: 24px; }
  .institutional-strip { grid-template-columns: 1fr; justify-items: center; gap: 30px; }
  .institutional-strip a { grid-column: auto; }
  .institutional-strip > img:first-child { width: 125px; }
  .institutional-strip > img:nth-child(2) { width: 250px; }
  .institutional-strip a img { width: 260px; }
  .related-panel > a { grid-template-columns: 80px 1fr; }
  .field-row { grid-template-columns: 1fr; }
  .site-footer { grid-template-columns: 1fr; text-align: center; justify-items: center; }
  .footer-brand { justify-self: center; }
  .site-footer nav { grid-column: auto; }
}

@media (prefers-reduced-motion: reduce) {
  html { scroll-behavior: auto; }
  *, *::before, *::after { animation-duration: .01ms !important; animation-iteration-count: 1 !important; transition-duration: .01ms !important; }
  .js [data-reveal] { opacity: 1; transform: none; }
}